Direct and indirect effects of climate change-amplified pulse heat stress events on coral reef fish communities.

Climate change-amplified temperature anomalies pose an imminent threat to coral reef ecosystems. While much focus has been placed on the effects of heat stress on scleractinian corals-including bleaching, mortality, and loss of reef structural complexity-and many studies have documented changes to reef fish communities arising indirectly from shifts in benthic composition, the direct impacts of heat stress on reef fish are much less well understood. Here, we quantify the direct and indirect effects of heat stress on reef fishes, using underwater visual censuses of coral reef fish communities conducted before, during, and after the 2015-2016 El Niño-induced global coral bleaching event.