Extreme armour in the world's oldest ankylosaur.

The armoured ankylosaurian dinosaurs are best known from Late Cretaceous Northern Hemisphere ecosystems, but their early evolution in the Early-Middle Jurassic is shrouded in mystery due to a poor fossil record. Spicomellus afer was suggested to be the world's oldest ankylosaur and the first from Africa, but was based on only a single partial rib from the Middle Jurassic of Morocco. Here we describe a new, much more complete specimen that confirms the ankylosaurian affinities of Spicomellus, and demonstrates that it has uniquely elaborate dermal armour unlike that of any other vertebrate, extant or extinct.

The world's oldest cerapodan ornithischian dinosaur from the Middle Jurassic of Morocco.

The cerapodan dinosaurs were an ornithischian clade that achieved a global distribution in the Cretaceous Period. The ichnological record suggests that these dinosaurs had evolved by the Middle Jurassic, but only a single cerapodan body fossil, an isolated femur from the Callovian of the UK, is known from this interval. In order to elucidate the early stages of cerapodan evolution and help to resolve the many phylogenetic inconsistencies in the clade, new specimens, particularly from historically undersampled localities, are needed.

New Middle to ?Late Jurassic dinosaur tracksites in the Central High Atlas Mountains, Morocco.

Besides bones, fossil tracks and trackways are important sources of knowledge about dinosaur palaeobiology. Here, we report three new tracksites from two different synclines in the Imilchil area, Central High Atlas, Morocco. The tracks and trackways are preserved in fluvial deposits in different levels of the Isli Formation (Early Bathonian-?Upper Jurassic), and contain impressions made by sauropods, theropods and ornithopods, as well as tracks that might represent bird-like non-avian theropod dinosaurs.

A second peirosaurid crocodyliform from the Mid-Cretaceous Kem Kem Group of Morocco and the diversity of Gondwanan notosuchians outside South America.

Notosuchians are an extinct clade of terrestrial crocodyliforms with a particularly rich record in the late Early to Late Cretaceous (approx. 130-66 Ma) of Gondwana. Although much of this diversity comes from South America, Africa and Indo-Madagascar have also yielded numerous notosuchian remains.

Bizarre dermal armour suggests the first African ankylosaur.

Ankylosauria is a diverse clade of armoured dinosaurs whose members were important constituents of many Cretaceous faunas. Phylogenetic analyses imply that the clade diverged from its sister taxon, Stegosauria, during the late Early Jurassic, but the fossil records of both clades are sparse until the Late Jurassic (~150 million years ago). Moreover, Ankylosauria is almost entirely restricted to former Laurasian continents, with only a single valid Gondwanan taxon.